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Dicot stem
Monocot stem
Broad pith
Weak stem
Monocot fiber sheaths
Ordinary cortex
Aerenchyma hypodermis
Aerenchyma cortex
Aerenchyma cortex 2
Stem endodermis
Palisade cortex
Cortical bundle
Capped cortical bundles
Collapsible cortex
Perimedullary fibers
Conjunctive tissue, paren.
Torn pith
Hollow pith
Medullary bundles
Typical dicot bundle
Vascular ring
Typical monocot bundle
Amphivasal bundle
Corn vascular bundle
Clintonia bundles
Protoxylem
Metaxylem
Metaxylem parenchyma
Metaxylem fibers
Internal phloem
Internal phloem, mag
Developing metaxylem
Primary phloem
Phloem fiber cap
Developing fibers

Fig. 11.4-3. Transverse section of ivy stem (Hedera helix). In some plants, the pith seems to play a minor role in storing water or starch or crystals, but in general, the pith of most stems is so small that it has too little volume to be a significant storage space. In a number of species, the pith seems to have no function whatsoever – its growth does not keep up with that of the surrounding vascular tissues, cortex and epidermis, so the pith is torn apart as internodes expand. This has occurred in this region of ivy stem, with the arrows indicating the many areas where pith cells have simply been torn apart. This is easy to see for yourself – just cut a young stem longitudinally and often you will find a hollow chamber rather than an intact pith.