Up Dicot stem Monocot stem Broad pith Weak stem Monocot fiber sheaths Ordinary cortex Aerenchyma hypodermis Aerenchyma cortex Aerenchyma cortex 2 Stem endodermis Palisade cortex Cortical bundle Capped cortical bundles Collapsible cortex Perimedullary fibers Conjunctive tissue, paren. Torn pith Hollow pith Medullary bundles Typical dicot bundle Vascular ring Typical monocot bundle Amphivasal bundle Corn vascular bundle Clintonia bundles Protoxylem Metaxylem Metaxylem parenchyma Metaxylem fibers Internal phloem Internal phloem, mag Developing metaxylem Primary phloem Phloem fiber cap Developing fibers
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Fig.
11.3-4.
Transverse section of stem cortex in sweetflag (Acorus). The cortex of
sweetflag becomes several millimeters thick, and this area is much closer to the
center of the stem than that of the previous micrograph. The intercellular
spaces here are even larger, and it is obvious that the “walls” of each
space consist of just a single layer of cortex cells: each cell is exposed to an
intercellular space on several sides. These spaces are probably schizogenous –
the young, developing tissue would have been compact but then at some point
during enlargement, the cells pulled apart from each other. We
know very little about the underlying mechanism – were the middle lamellas
torn apart in certain areas or were they digested enzymatically by the cells?
What mechanism determined which areas of middle lamella would be ruptured and
which would remain intact?
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